Angelica and hemp agrimony - seething with insect life in late summer

 Angelica sylvestris, a member of the Apiaceae (carrot family) is surely one of the best umbellifers for supplying insects with pollen and nectar. It thrives in moist soils places, often in dappled sunlight along woodland edges. The pictures below are of a greenbottle, a Darwin wasp (ichneumon) and social paper wasps feeding on nectar from its numerous tiny florets. The nectar is secreted in small amounts on each individual flower, so is easily accessible to short-tongued insects, without them needing to expend much energy; all they need do is wander across the umbel, incidentally pollinating the flowers as they feed.

Hemp agrimony Eupatorium cannabinum, a tall, pink-flowered member of the daisy family (Asteraceae) seems to be flowering spectacularly well along the Durham coast near Nose's Point this summer, in the company of meadowsweet and creeping thistle. Soldier beetles (last photo) are amongst the vast number of insects that visit its flowers, in the company of bees, butterflies and many species of fly.

A hay meadow going to seed

Monday's Guardian Country Diary is an account of a wonderful hay meadow at the eastern end of Hawthorn dene, a Durham Wildlife Trust nature reserve on the Durham coast. 

Most of the Pennine hay meadows are cut at around the third week of July, which allows the farmer to take a decent hay crop but also allows key meadow wild flowers - like hay rattle - to set seed. This meadow is managed entirely for its glorious native flora and so was still uncut when we visited in late August. There were still plenty of species in flower, including knapweed, meadowsweet, field scabious, devils bit scabious and hemp agrimony, but all the plants here were also producing masses of seed. 

This view was taken looking northwards .......

.......... this one is from the coast side, looking towards the wooded dene ...

.......... and this is the view to the south. In spring this field holds thousands of cowslips, together with early purple orchids and wood cranesbill. In summer meadow and bloody cranesbill are prominent. When I walked across here I counted about fifty species without even bending down to have a close look - I suspect that there are least twice as many here. 

Although it's cut very late, it still must be cut and a hay crop taken, because there is a constant rain of seeds in autumn from the wooded dene and without the mower it would soon become ash, sycamore and hawthorn shrub - last autumn's tree seedlings were already well established. 

Field scabious flowers produce these attractive hemispheres of bristly seeds after pollination

The late flowering hemp agrimony is a great attraction for butterflies like this comma.

Devil's bit scabious, one of the most attractive late summer wild flowers in this limestone grassland, and so called because the stumpy root looks like Old Nick himself has taken a bite out of it.

Although vast quantities of seed are produced by the plants here only a small minority germinate in any single year and become mature plants, simply because there is so much competition in a dense sward with so many species. The best germination sites are mole hills like this, which provide a perfect vacant seed bed for any seed fortunate enough to land on them. Every hay meadow needs mole hills.

...... and now for the good news.....

If you let it, taking an interest in natural history could be very depressing because so much that is written about our flora and fauna in the press seems to be unrelenting bad news. The abysmal weather in the early part of this summer  produced also sorts of hyperbolic headlines of the disastrous/cataclysmic/catastrophic kind about its likely impact on birds/butterflies/insects and almost everything else in the natural world - and it's undoubtedly true that it has been a bad summer for all of these (although it's important to bear in mind that over last few thousand years there have been periods of far more adverse climatic conditions that our wildlife experienced and rebounded from: like investments, rates of growth in populations of living organisms can go down as well as up, all the time). The natural world isn't a natural history museum, it's a living, constantly evolving interplay between organisms.

Anyway, last weekend we happened to notice a positive effect of the wet early part of the summer, when we took a walk along the cliff tops between Dawdon and Hawthorn Dene on the Durham Coast Heritage Path. The summer wild flowers there seem to be flowering more spectacularly than I can ever remember. I have a theory as to why this might be so, and it's all down to the rain. 

The cliffs are porous magnesian limestone which is free-draining. In a 'normal' summer the surface soil dries out quickly and the plants suffer from moisture stress, so they flower quickly while they have the chance, when they are quite small. This year, with plenty of water, the well-watered vegetation is unusually lush and as a result the flowers are truly spectacular. So, without further ado, here are some examples....

Drifts of meadowsweet in the gullies on the cliffs and ....

.... forming a backdrop for devil's bit scabious which .....

...... includes some exceptionally fine examples ..... although this one looks a tad unusual - very short bracts under the inflorescence and it looks like it might be male-sterile, judging by the tiny stamens.

... along with field scabious .....

... flowering in profusion in the meadows at the end of Hawthorn Dene ...

... where there are some fine specimens of giant bellflower.

I suspect that alkaline soil might be the reason for the very blue hue of the tufted vetch on the cliffs but .... 

... this unusually pale betony plant (normal flower behind) is probably a genetic mutation.

Bloody crane'sbill always puts on a good show ....

.... and this year centaury has done well too, with some very floriferous plants ...

This is hemp agrimony .....a major constituent of the taller vegetation while .....

.... hoary plantain is blooming down amongst the grasses.

Plenty of knapweed Centaurea nigra, alongside the ....

.... greater knapweed Centaurea scabiosa.

Drifts of marjoram aka oregano (crush the leaves and sniff for a tantalising hint of Mediterranean cuisine!)....

Spear thistle already running to seed ....

.... likewise agrimony, producing its little bell-shaped hooked fruits ....

Yellow loosestrife ....

.... and some wonderfully robust specimens of yellow-wort, with its strange glaucous, stalkless leaves.

In dry summers yellow-wort, which is an annual, is often a stunted plant and only produces a few flowers - but this year some plants have responded to the rain with a wonderful display...

... and the North Sea provides a fine backdrop for the floral spectacle ....

..... and for haymaking ....